Last updated Feb 07, 2026
*Undergraduate mentee; †Co-first authors
Pan, V. S., Rothstein, P. E.*, Gilbert, K. J. 2026. The Frequency of Population and Masting Fluctuations Is Increasing Globally. bioRxiv. [link] [PDF]
Karban, R., Pepi, A. A., Grof-Tisza, P., Pan, V. S., Loeb, G., Huntzinger, M. and Holyoak, M. 2026. Precipitation Drives the Abundance and Distribution of Arctia virginalis: A 40-Year Study. BioScience. In press: biaf203. [link] [PDF]
Pan, V. S., Gilbert, K. J., Wetzel, W. C. 2025. Mean Plant Toxicity Modulates the Effects of Plant Defense Variability. Ecology. 106:e70012. [link][PDF]
Pan, V. S., Ghosh, E., Ode, P. J., Wetzel, W. C., Gilbert, K. J., Pearse, I. S. 2025. Large Differences in Herbivore Performance Emerge from Simple Herbivore Behaviors and Fine-Scale Spatial Heterogeneity in Phytochemistry. Ecology Letters. 28:e70044. [link][PDF]
Pan, V. S., Wetzel, W. C. 2024. Neutrality in Plant–Herbivore Interactions. Proceedings of the Royal Society B: Biological Sciences. 291:20232687. [link][PDF]
Robinson, M.L., Hahn, P.G., Inouye, B.D., Underwood, N., Whitehead, S.R., Abbott, K.C., Bruna, E.M., Cacho, N.I., Dyer, L.A., Abdala-Roberts, L., Allen, W.J., Andrade, J.F., Angulo, D.F., Anjos, D., Anstett, D.N., Bagchi, R., Bagchi, S., Barbosa, M., Barrett, S., Baskett, C.A., Ben-Simchon, E., Bloodworth, K.J., Bronstein, J.L., Buckley, Y.M., Burghardt, K.T., Bustos-Segura, C., Calixto, E.S., Carvalho, R.L., Castagneyrol, B., Chiuffo, M.C., Cinoğlu, D., Cinto Mejía, E., Cock, M.C., Cogni, R., Cope, O.L., Cornelissen, T., Cortez, D.R., Crowder, D.W., Dallstream, C., Dáttilo, W., Davis, J.K., Dimarco, R.D., Dole, H.E., Egbon, I.N., Eisenring, M., Ejomah, A., Elderd, B.D., Endara, M.-J., Eubanks, M.D., Everingham, S.E., Farah, K.N., Farias, R.P., Fernandes, A.P., Fernandes, G.W., Ferrante, M., Finn, A., Florjancic, G.A., Forister, M.L., Fox, Q.N., Frago, E., França, F.M., Getman-Pickering, A.S., Getman-Pickering, Z., Gianoli, E., Gooden, B., Gossner, M.M., Greig, K.A., Gripenberg, S., Groenteman, R., Grof-Tisza, P., Haack, N., Hahn, L., Haq, S.M., Helms, A.M., Hennecke, J., Hermann, S.L., Holeski, L.M., Holm, S., Hutchinson, M.C., Jackson, E.E., Kagiya, S., Kalske, A., Kalwajtys, M., Karban, R., Kariyat, R., Keasar, T., Kersch-Becker, M.F., Kharouba, H.M., Kim, T.N., Kimuyu, D.M., Kluse, J., Koerner, S.E., Komatsu, K.J., Krishnan, S., Laihonen, M., Lamelas-López, L., LaScaleia, M.C., Lecomte, N., Lehn, C.R., Li, X., Lindroth, R.L., LoPresti, E.F., Losada, M., Louthan, A.M., Luizzi, V.J., Lynch, S.C., Lynn, J.S., Lyon, N.J., Maia, L.F., Maia, R.A., Mannall, T.L., Martin, B.S., Massad, T.J., McCall, A.C., McGurrin, K., Merwin, A.C., Mijango-Ramos, Z., Mills, C.H., Moles, A.T., Moore, C.M., Moreira, X., Morrison, C.R., Moshobane, M.C., Muola, A., Nakadai, R., Nakajima, K., Novais, S., Ogbebor, C.O., Ohsaki, H., Pan, V.S., Pardikes, N.A., Pareja, M., Parthasarathy, N., Pawar, R.R., Paynter, Q., Pearse, I.S., Penczykowski, R.M., Pepi, A.A., Pereira, C.C., Phartyal, S.S., Piper, F.I., Poveda, K., Pringle, E.G., Puy, J., Quijano, T., Quintero, C., Rasmann, S., Rosche, C., Rosenheim, L.Y., Rosenheim, J.A., Runyon, J.B., Sadeh, A., Sakata, Y., Salcido, D.M., Salgado-Luarte, C., Santos, B.A., Sapir, Y., Sasal, Y., Sato, Y., Sawant, M., Schroeder, H., Schumann, I., Segoli, M., Segre, H., Shelef, O., Shinohara, N., Singh, R.P., Smith, D.S., Sobral, M., Stotz, G.C., Tack, A.J.M., Tayal, M., Tooker, J.F., Torrico-Bazoberry, D., Tougeron, K., Trowbridge, A.M., Utsumi, S., Uyi, O., Vaca-Uribe, J.L., Valtonen, A., van Dijk, L.J.A., Vandvik, V., Villellas, J., Waller, L.P., Weber, M.G., Yamawo, A., Yim, S., Zarnetske, P.L., Zehr, L.N., Zhong, Z., Wetzel, W.C., 2023. Plant size, latitude, and phylogeny explain within-population variability in herbivory. Science. 382:679–683. [link][PDF]
Pepi A, Pan, V. S., Grof-Tisza, P., Holyoak, M., Balldman, A., Laws-McNeil, A., Mase, V., Moseley, C., Karban, R. 2023. Spatial habitat heterogeneity influences host-pathogen dynamics in a patchy population of Ranchman’s tiger moth. Ecology. 104:e4144. [link][PDF]
LoPresti, E. F., Pan, V. S., Girvin, C., Barber, G., Jaeger, S., Orrock, J. 2023. Sandy seeds: armor or invisibility cloak? Mucilage-bound sand physically protects seeds from rodents and invertebrates. Journal of Ecology. 111:540–551. [link] [PDF]
Pan, V. S., A. Pepi, E. F. LoPresti, R. Karban. 2023. The Consequence of Leaf Life Span to Virus Infection of Herbivorous Insects. Oecologia. 201:449–459. [link] [PDF]
Pepi, A., Pan, V. S., Rutkowski, D., Mase, V. Karban, R. 2022. Influence of delayed density and ultraviolet radiation on caterpillar baculovirus infection and mortality. Journal of Animal Ecology. 91:2192–2202. [link] [PDF]
Orrock J. L., Guiden, P. W., Pan, V. S., Karban, R. 2022. Plant induced defenses that promote cannibalism reduce herbivory as effectively as highly pathogenic herbivore pathogens. Oecologia. 199:397–405. [link] [PDF]
Pan, V. S., Girvin, C., LoPresti, E. F. 2022. Anchorage by seed mucilage prevents seed dislodgement in high surface flow: a mechanistic investigation. Annals of Botany. 129:817–830. [link] [PDF]
Pan, V. S., Marshall, M., Karban, R., Goidell, J., Weber, M. G., LoPresti, E. F. 2021. Mucilage binding to ground protects seeds of many plants from harvester ants: A functional investigation. Functional Ecology. 35:2448–2460. [link] [PDF]
Pan, V. S., Pepi, A., Goidell, J., Karban, R. 2021. Retention of green leaves not brown leaves increases spring cynipid diversity on large valley oaks. Arthropod-Plant Interactions. 15:353–362. [link] [PDF]
Grof-Tisza, P., Karban, R., Pan, V. S., Blande, J. D. 2020. Assessing plant-to-plant communication and induced resistance in sagebrush using the sagebrush specialist Trirhabda pilosa. Arthropod-Plant Interactions. 14:327–332. [link] [PDF]
Nguyen, H. D. D., Pan, V. S., Pham, C., Valdez, R., Doan, K., Nansen, C. 2020. Night-based hyperspectral imaging to study association of horticultural crop leaf reflectance and nutrient status. Computers and Electronics in Agriculture. 173:105458. [link] [PDF]
LoPresti, E. F., Pan, V. S., Goidell, J., Weber, M. G., Karban, R. 2019. Mucilage-bound sand reduces seed predation by ants but not by reducing apparency: a field test of 53 plant species. Ecology. 100:e02809. [link] [PDF]
LoPresti, E. F., Pan, V. S., Goidell, J., Weber, M. G., Karban, R. 2019. Mucilage-Bound Sand Reduces Seed Predation by Ants But Not by Reducing Apparency: A Field Test of 53 Plant Species. The Bulletin of the Ecological Society of America. 100:e01596. [link] [PDF]